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In flowering plants, gibberellin (GA) regulates the formation of the outer wall on pollen and is therefore essential for pollen development1,2,3. The outermost pollen wall, which is known as the exine or exospore, consists largely of sporopollenin, an organic polymer that is extremely resistant to degradation and is so durable that it has been found in microfossils that are over 500 million years
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