Crustle is an orange Pokémon resembling an insect or crustacean. Its most distinguishing feature is its enormous shell, which resembles a cut-away block of sediment with multiple layers and strata visible. The tips of its pincers and six feet are dark gray. It has two glaring yellow eyes. Crustle possesses a straight tail that is concealed underneath its shell.
It is very strong, to the point that it can carry heavy rocks for a long time, even through arid lands where it resides. It fights members of its species over territory in battles which can become very vicious. If its shell breaks during such a fight, it loses. When Crustle's shell breaks, it will become weaker and unsure of itself. Crustle's biggest strength is its thick claws; being strong enough to break even Rhyperior's sturdy carapace.
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Crustle is based on a hermit crab. Its shell resembles a block of stratum rather than an actual shell, though the design also seems to resemble ornately-colored hermit crab shells.