Photography by Bj�rn R�rslett/NN
The compilation of species will continue to be updated at irregular intervals. All species listed here have been documented, and links are added whenever I can find spare time for updating. These images are made for illustrative purposes, not as artistic statements per se. However, there are lots of food for thought in the convoluted ways Nature expresses itself, so for once the artist can step backand let the subjects speak for themselves. "Das Ding an Sich" to paraphrase Kant, or Eigenvalue of Nature. If you are unfamiliar with the botany, just select any species indicated as having a "strong" response to learn how this looks. Potentilla anserina may constitute an introductory example. In case you wonder about the family and species selection, these are plants readily available to me. However, not all species have the typical bull's-eye UV pattern, which may be confined to symmetrical flowers. Nevertheless flowers may exhibit a virtually endless variety of spectral signatures. Just take a look at this modest plant, Glechoma hederacea, to get an impression of the near bewildering spectral diversity that exists. The UV range of the spectrum has no predefined colours, so we are free to assign any colour we like. There are no rules set in stone for this at all. I set out my UV work using film, so in principle would like the "film UV colours" (blue with UV absorbance often shown in reddish hues) to prevail. Different spectral response of the UV bandpass filters applied in these years, in conjunction with changing behaviour of the digital cameras and their RAW-file converters, have made this aim impossible to uphold. Thus, don't compare colours, compare patterns. I'm currently writing a tutorial on how the UV "colours" are created and how they should be interpreted, but for the time being, just enjoy the images like the pollinators enjoy their targeted flowers. UV fluorescence may be a common trait to most flowers, but might be of temporary occurrence for parts of the flower. Anthers, style, and pollen grains occasionally are seen to fluoresce. Strong fluorescence has been noted from nectar glands (Angelica sylvestris) and several other species. Some species show fluorescence of the non-fertilised stigmas, but this trait is difficult to document with my normal technical approach. Fluorescence from outside of the bracts is exhibited by some species. As far as the photography is concerned, the main issue with flower fluorescence is its transient behaviour. It may be present, but the flowers collected for photography don't appear to fluoresce simply because the floral development is in the "wrong" stage. With fluorescent pollen grains, their size often are at or below the detection limit unless quite high magnification is employed, thus calling for a true photomacropgraphic approach. The fluorescing pollen of Mirabilis jalapa (Four o'Clock) has been documented using this method. In case you are curious as to why the species might have these patterns, read this to learn more. UV-absorbing substances (flavonyl glucosides) are instrumental in bringing about the fascinating pollinating guide patterns. UV marks on flowers are but a logical extension of the visual pollinating clues provided by evolution in Nature. When a flower is stated to lack UV marks, it simply means the signals emitted are directed towards pollinators which can "see" in alternate spectral bands, outside the UV range. If the flower absorbs UV all over the floral parts, it may appear visually in a "UV-complementary" colour even to pollinators capable of seeing in UV. We can only speculate as to the rendition of that complementary colour, but if say the insect is modelled as seeing UV as "blue", blue as "green", and green as "red", then the UV complementary would be yellow. Thus, a UV-absorbing yellow flower still would come across as "yellow" even for an insect (or so it might seem, but who are we to know such things anyway). A final word of warning is that UV photography as such has lots of technical pitfalls and issues. You have to make absolutely sure your camera really records UV patterns, and in order to ensure this, careful testing of your setup is necessary. This has become even more important in the digital era, as outlined elsewhere in my UV/IR tutorial. Read in particular about the issues with IR contamination described there. |
Plant/ Family |
Flower Colour | Ultraviolet Features
(Strong, intermediate etc. refer to a UV-visible bull's-eye pattern of the flower. Other patterns may appear as duly noted) |
Amaryllidaceae | ||
Narcissus pseudonarcissus L. | Bright-deep yellow | Not a bull's eye pattern, but the flower appears flamed in UV light with alternating high- and low-UV reflecting stripes |
Apiaceae | ||
Angelica sylvestris L. | White | Low UV reflectance of entire flower, but very strong UV fluorescence of the nectar glands |
Carum carvi L. | Dullish white or reddish | � |
Apocynaceae | ||
Vinca minor L. | Light violet | Strong UV reflectance, but a bull's eye pattern is not very prominently developed. High reflectance in near IR, though |
Asteraceae |
||
Achillea millefolium L. | Dullish white | Very weak |
Anthemis tinctoria L. | Dark yellow | Intermediate - strong, some UV fluorescence |
Arnica angustfifolia Vahl | Dark yellow | Very strong bull's eye pattern |
Aster tripolium L. | Light violet | Intermediate, coincides with visual pattern |
Bidens sp. | Deep yellow | Big, very prominent UV marks |
Calendula officinalis L. | Orange | Strong bull's eye |
Centaurea jacea L. | Purple | Intermediate - strong |
Chrysanthemum segetum L. | Bright yellow | None (all the flower head absorbs UV) |
Cicerbita alpina (L.)Wallr. | Light violet | Intermediate |
Cichorium intybus L. | Light blue | Intermediate - strong UV marking |
� | � | � |
Crepis biennis L. | Bright yellow | Strong, but not well delineated |
Doronicum orientale Hoffm. | Deep yellow | Very strong bull's eye pattern. UV fluorescence. |
Hieracium aurantiacum L. | Reddish | Unusual, chequered pattern |
Hieracium subg. Pilosella | Yellow | Variegated UV pattern |
Hieracium sect. Prenathaea | Yellow | Strong |
Hieracium sect. Sylvatica | Yellow | Strong |
Lapsana communis L. | Yellow | Strong |
Leucanthemum vulgare Lam. | White petals, yellow centre disc | Weak, some zonation across central floral disc, low UV reflectance of outer flower areas |
Leontodon autumnale L. | Yellow | Strong |
Mycelia muralis (L.)Dum. | Light yellow | Intermediate - strong |
Petasites hybridus (L.)Gaertner, Meyer & Scherb. | Very light pink, rarely light yellow | Strong UV fluorescence |
Rudbeckia hirta L. | Bright yellow | Strong. Very large and prominent UV marks |
Senecio aquaticus Hill. | Bright yellow | Very strong, entire centre absorbs UV |
Senecio jacobea L. | Deep yellow | Very strong |
Senecio viscosus L. | Light yellow | Ray flowers strongly reflect UV |
Sonchus arvensis L. | Deep yellow | Strong bull's eye pattern, but UV delineation against the flower periphery is blurred (a common trait to UV marks of many composite flowers, by the way) |
Sonchus oleraceus L. | Light yellow | Strong |
Tanacetum vulgare L. | Deep yellow | Negligible. Some zonation across the floral discs on the onset of flowering |
Taraxum vulgare Web. | Yellow | Usually extremely pronounced, but quite variable extent of bull's-eye pattern. Dandelions are really a cluster of related, apomictic microspecies. This helps explain why the UV markings may vary so much |
Tragopogon pratensis L. ssp. minor (Mill.)Hartm. | Deep yellow | No typical bull's eye pattern, but strongly UV-reflective bracts contribute to form a two-zoned flower head |
Tripleurospermum maritium (L.)W.D.J. Koch | White | Strong bull's eye |
Tussilago farfara L. | Yellow | Strong bull's-eye pattern, but variable UV reflectance of outer petals |
Brassicaceae |
||
Alliaria petiolata (Bib.)Cavara & Grande | White | Intermediate. Anthers and pollen absorb strongly in UV |
Barbarea vulgaris (L.)R.Br. | Yellow | Strong |
Berteroa incana (L.)DC. | White | Not a typical bull's eye pattern in its small flowers. However, there are floral buds and stigma with a prominent UV fluorescence |
Brassica nigra (L.) Koch | Yellow | Strong bull's eye pattern |
Bunias orientalis L. | Light yellow | No bull's eye pattern, but quite high UV reflectance of petals |
Erysimum hieracifolium L. | Yellow | Petals darkish under UV light, buds and non-fertilised flowers show strong UV fluorescence |
Hesperis matronalis L. | Pink, mauve or purple | Intermediate, UV fluorescence |
Rorippa sylvestris (L.)Bess. | Yellow | Intermediate - strong |
Sinapis arvensis L. | Yellow | Very strong bull's-eye pattern, young buds absorb UV, too |
Campanulaceae |
||
Campanula glomerata L. | Dark purple | Intermediate to strong two-zoned pattern, very high UV reflectance of outer floral parts |
Campanula latifolia L. | White or blue | � |
Campanula rapunculoides L. | Light violet | Intermediate to strong UV marking |
Campanula persicifolia L. | Deep or sky blue | Very high UV reflectance of corolla, high UV absorption from stigma and anthers |
Campanula rotundifolia L. | Light to sky blue | Intermediate, UV marks mainly on pistils and stigma |
Campanula trachelium L. | Bright-dullish blue, with greyish tint | Very high UV reflectance of corolla and buds, stigma absorbs UV, nectaria fluoresce in UV |
Jasione montana L. | Bright blue | Flower heads with a mosaic of UV-absorbing or reflecting areas |
Lobelia dortmanna L. | White or pinkish purple | � |
Caprifoliaceae |
||
Lonicera perclymemum L. | Creamy white | Weak |
Caryophyllaceae | ||
Cerastium arvense L. | White | Strong |
Dianthus deltoides L. | Pinkish or light purple | Strong, high IR reflectance, UV fluorescence |
Lychnis viscaria L. | Bright purple | Strong, high IR reflectance |
Saponaria officinalis L. | Light pink | Petals and anthers absorb UV, the border of petals do reflect UV |
Silene armeria L. | Bright purple | Intermediate - high UV reflectance, depending on floral stage |
Silene dioica (L.)Clairv. | Bright purple or red | Strong UV pattern, high IR reflectance |
Silene uniflora Roth | White | Intermediate |
Convolvulaceae |
||
Calystegia sepium (L.)R.Br. | Pinkish or white | Intermediate - strong UV pattern |
Convolvulus arvensis L. | White and pink | Intermediate, reflects also strongly in IR |
Cuscuta europaea L. ssp. halophyta (Fr.)Hartm. | Pinkish white | Whole corolla absorbs UV strongly, some UV fluorescence from young flowers |
Crassulaceae |
||
Sedum acre L. | Yellow | Intermediate - strong |
Sedum reflexum | � | � |
Sedum telephium L. | Light yellowish | None |
Dipsacaceae | ||
Knautia arvensis (L.)Coult. | Purplish-violet | Intermediate - strong, unusual chequered UV pattern |
Droseraceae | ||
Drosera longifolia L. (flowers) | White | Petals, anthers, and stigmas absorb UV, nectaria have strong UV fluorescence |
Drosera longifolia L. (leaves) | (yellowish green, red tentacles) |
Leaf blade absorbs UV, viscous fluid drops on insect-catching tentacles have strong UV fluorescence |
Ericaceae |
||
Andromeda polifolia L. | Pink red | � |
Calluna vulgaris L. | Greyish purple | � |
Erica tetralix L. | Pinkish red | � |
Loiseleuria procumbens (L.)Dum | Pinkish white | Intermediate - strong |
Fabaceae |
||
Lathyrus linifolius (Reichard)B�ssler | Variegated purple - greyish violet | No bull's eye pattern, but high UV reflectance of corolla |
Lathyrus odoratus L. | Pink | No bull's eye pattern, but significant UV reflectance of corolla |
Lathyrus pratensis L | Yellow | No bull's eye pattern, but high UV reflectance of corolla |
Lotus corniculatus L. | Yellow or Reddish tinged | Indistinct, variegated stripes on corolla |
Vicia sepium L. | Mauve or light pink | Very weak UV signs, but the corolla does reflect UV and additonally shows fluorescence |
Gentaniaceae | ||
Centaurium littorale (Turner) Gilmour | Pink or light purple | Anthers and stigma strongly absorb UV |
Geraniaceae |
||
Geranium pratense L. | Bluish or purple, often with a greyish tinge | Variable-sized central UV absorbing spot, combined with UV-absorbing rays radiating outwards. IR reflectance is quite high, too |
Geranium sanguineum L. | Scarlet or deep purplish red | Strong, in combination with a radiating pattern on the petals. Strong IR reflectance |
Geranium robertianum L. | Light purple | Strong bull's-eye marking |
Geranium sylvaticum L. | Purplish red | Very strong bull's-eye pattern. Strong reflectance in near-IR |
Hypericaceae | ||
Hypericum maculatum Cr. | Deep yellow | Intermediate - strong |
Hypericum perforatum L. | Deep yellow | Intermediate - strong |
Lamiaceae |
||
Glechoma hederacea L. | Bluish violet or purple | No typical bull's eye pattern, but variegated stripes on petals from UV reflectance. Also distinguished by anthers which both reflect UV and show strong fluorescence under UV light. In addition, whole flower reflects strongly in the near-IR band. |
Stachys officinalis (L.)Trev. | Purple | Large areas of the corolla reflect UV strongly |
Liliaceae | ||
Gagea lutea (L.)Ker-Gawler | Yellow | Strong |
Gagea minima (L.)Ker-Gawler | Yellow | Strong |
Lilium bulbiferum L. | Red | Intermediate, some UV fluorescence |
Lythraceae | ||
Lythrum salicaria L. | Purple | Intermediate |
Nyctaginaceae | ||
Mirabilis jalapa L. | Yellow | Moderate UV reflectance of perianth, but significant UV fluorescence of corolla and pollen |
Nymphaeaceae |
||
Nuphar lutea (L.)Sibth. & Sm. | Yellow | � |
Nuphar
lutea x pumila (N. spenneriana) |
Yellow | Strong, unusual circular pattern |
Nuphar pumila (Timm)DC. | Yellow | � |
Nymphaea alba L. | White | Intermediate |
Onagraceae | ||
Epilobium angustifolium L. | Purplish red | Weak |
Epilobium montanum L. | Light red | None |
Epilobium watsonii | Light red | � |
Oenothera biennis L. | Light yellow | Prominent and stunningly beautiful UV markings |
Oxalidaceae | ||
Oxalis fontana Bunge | Light yellow | Strong |
Papaveraceae | ||
Eschscholzia californica Cham. | Bright yellow with orange marks on petals | Very large UV marks, not coinciding with visual pattern |
Plantaginaceae | ||
Plantago media L. | Light pink | Very weak within the floral parts, some UV reflectance from upper part of stems and spikes |
Primulaceae |
||
Lysimachia nummularia L. | Yellow | Weak |
Lysimachia punctata L. | Yellow | None |
Ranunculaceae |
||
Anemone blanda Schott & Kotsky | Purplish blue | Strong UV pattern, high IR reflectance as well |
Anemone nemorosa L. | White | No bull's eye pattern discernible. Quite strong reflectance in near-IR |
Anemone ranunculoides L. | Deep yellow | Strong, indistinct secondary markings on petals |
Caltha palustris L. | Deep yellow | Strong, very large central zone with UV mark. Complex UV fluorescence patterns |
Eranthis hyemalis (L.)Salisb. | Yellow | Intermediate - strong |
Hepatica nobilis Schreb. | Light blue | Very strong UV reflectance (and in near-IR as well) |
Ranunculus acris L. | Yellow | Very prominent bull's-eye pattern, despite only moderate UV reflectance of the flower. A weak two-zoned pattern is present even in near-IR |
Ranunculus auricomis L. | Yellow | Weak, non-descript UV pattern |
Ranunculus ficaria L. | Bright yellow | Very strong bull's pattern in UV, also a similar pattern (but much weaker) found in near-IR |
Ranunculus repens L. | Yellow | Weak - intermediate |
Ranunculus reptans L. | Yellow | Strong, base of petals |
Rosaceae |
||
Agrimonia eupatoria L. | Yellow | High UV reflectance of petals, nectaria and styles absorb UV |
Comarum palustre L. | Dark red or purple | � |
Fragaria vesca L. | White | Weak, but detectable |
Fragaria vesca L. x viridis Duch. | White | Interestingly, the UV marks are intermediate to those of the parents |
Fragaria viridis Duch. | White | A bull's eye pattern is present, but not very strongly developed as the outer petals themselves reflect only moderately in UV. The bracts, however, are good UV reflectors |
Geum chilo�nse Balb. | Yellowish red | Moderate - strong |
Geum urbanum L. | Yellow | Moderate - strong |
Potentilla anserina L. | Dark yellow | Extremely strong pattern, making this an ideal test object for UV photography |
Potentilla atrosanguinea Lodd. | Reddish with darker centre | Strong, but UV marks are located slightly different from visual clues |
Potentilla erecta (L.)R�usch. | Yellow | Extremely strong, excellent test object. UV fluorescence |
Potentilla fruticosa L. | Bright yellow | None. Petals with negligible UV reflectance |
Potentilla intermedia L. | Deep yellow | Strong, bracts and receptacle show UV fluorescence |
Potentilla norvegica L. | Light yellow | Intermediate - strong |
Potentilla recta L. | Deep yellow | Very strong, prominent UV marks. UV fluorescence of pollen and nectaria |
Potentilla reptans L. | Deep yellow | Very strong and prominent UV marks |
Potentilla rupestris L. (Drymocallis rupestris) |
White | Weak or intermediate |
Rosa dumalis Bechst. | Pinkish red | Weak, but present |
Rosa majalis L. | Reddish | Weak to intermediate |
Scrophulariaceae |
||
Digitalis purpurea L. | Light red or creamy white | Intermediate - strong |
Melampyrum pratense L. | Yellowish | Weak |
Linaria vulgaris L. | Light yellow | None |
Verbascum nigrum L. | Yellow | Intermediate - strong |
Verbascum phlomoides L. | Yellow | Intermediate - strong |
Solanaceae |
||
Hyoscyamus niger L. | Dirty yellow and brown | Very strong bull's-eye pattern present, also strong IR reflectance |
Solanum dulcamara L. | Purple | Weak - intermediate, not a typical bull's-eye pattern |
Solanum niger L. | Whitish | � |
Violaceae |
||
Viola arvense x tricolor | White to yellow, some violet | Weak UV reflectance, no clear-cut pattern |
Viola odorata L. | Violet | None, but slight UV reflectance of petals |
Viola tricolor L. | Tricoloured white, yellow, and violet | Weak, more or less coinciding with the visual clues |
Note: Some species may have a flower response in the near-infrared (nIR) as well. Few occurrences of this are yet documented, but Ranunculus ficaria might serve as an example. I haven't yet decided whether or not IR aspects should be addressed in these studies. More test shots are needed to assess the potential of adding IR to the study. Since my UV work receives no external funding, I have limits as to the efforts and time which can be spent on documenting these flower features.
All images on these web pages are copyright Bj�rn R�rslett - NN/N�rfoto and cannot be used for any purpose, commercial or non-commercial, without express and written approval by the author.
Most uses will incur a negotiable fee. Please e-mail me for details.
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Last Update 24 October, 2006